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Sceptridium is a genus of seedless vascular plants in the family Ophioglossaceae,cite-ref-christenhusz-2011b-1-0[1] closely allied to (and often included as a subgenuscite-ref-2[2] of) the genus Botrychium (the moonworts and grapeferns). It is also closely related to the genus Botrypus (the rattlesnake fern, often treated as the subgenus Osmundopteris under Botrychium). Sceptridium species are commonly called the grape-ferns.

These plants are small with fleshy roots, and reproduce by spores shed into the air. They differ from the moonworts in having at least some sterile fronds (all fronds in Botrychium are spore-bearing), and in the fronds being bi- or tri-pinnate (Botrychium are single pinnate, or rarely bipinnate); and from Botrypus in being evergreen, or at least winter-green (Botrypus are deciduous) and having the non-spore-bearing part of the frond long-stalked (short-stalked in Botrypus).

• Phylogeny

• References

Phylogeny of Sceptridiumcite-ref-3[3]cite-ref-4[4]

Unassigned species:

• S. × argutum Sahashi

• S. australe (Brown) Lyon (parsley fern)cite-ref-22[22]cite-ref-23[23]

• S. biforme (Col.) Lyon (fine-leaved parsley fern)cite-ref-24[24]cite-ref-25[25]

• S. × elegans Sahashi

• S. javanicum Sahashi

• S. jenmanii (Underwood) Lyon (Alabama grape-fern)cite-ref-26[26]cite-ref-27[27]cite-ref-28[28]

• S. × longistipitatum Sahashi

• S. × pulchrum Sahashi

• S. rugulosum (Wagner) Skoda & Holub (ternate grape-fern, St. Lawrence grape fern, rugulose grape fern)cite-ref-mich-7-3[7]cite-ref-29[29]cite-ref-botrug-30-0[30]

• S. × silvicola (Sahashi) Ebihara

• S. subbifoliatum (Brack.) Lyon (island grape-fern)cite-ref-31[31]cite-ref-32[32]cite-ref-33[33]

• S. underwoodianum (Maxon) Lyoncite-ref-34[34]cite-ref-35[35]

cite-note-christenhusz-2011b-11. ↑ citerefchristenhuszzhangschneider2011Christenhusz, Maarten J. M.; Zhang, Xian-Chun; Schneider, Harald (2011). "A linear sequence of extant families and genera of lycophytes and ferns" (PDF). Phytotaxa. 19: 7–54. doi:10.11646/phytotaxa.19.1.2.

cite-note-22. ↑ subgenus Tropicos.org. Missouri Botanical Garden. 16 Jan 2012

cite-note-33. ↑ citerefnittaschuettpelzram-rez-barahonaiwasaki2022Nitta, Joel H.; Schuettpelz, Eric; Ramírez-Barahona, Santiago; Iwasaki, Wataru; et al. (2022). "An Open and Continuously Updated Fern Tree of Life". Frontiers in Plant Science. 13: 909768. doi:10.3389/fpls.2022.909768. PMC 9449725. PMID 36092417.

cite-note-44. ↑ "Tree viewer: interactive visualization of FTOL". FTOL v1.7.0 (GenBank release 261). 2024. Retrieved 1 May 2025.

cite-note-51. "PteridoPortal - Sceptridium lunarioides". www.pteridoportal.org. Retrieved 7 October 2020.

cite-note-62. B. schaffneri Tropicos.org. Missouri Botanical Garden. 04 Jan 2012

cite-note-mich-77. [1] MICHIGAN FLORA ONLINE. A. A. Reznicek, E. G. Voss, & B. S. Walters. February 2011. University of Michigan. Web. January 16, 2012

cite-note-83. citerefmesipuur-p-sheffersont-kull2009Mesipuu, Meeli; R. P. Shefferson; T. Kull (2009). "Weather and herbivores influence fertility in the endangered fern Botrychium multifidum (S.G. Gmel.) Rupr". Plant Ecology. 203 (1): 23–31. Bibcode:2009PlEco.203...23M. doi:10.1007/s11258-008-9501-3. S2CID 13072960.

cite-note-94. B. multifidum Tropicos.org. Missouri Botanical Garden. 26 Dec 2011

cite-note-105. B. multifidum Flora of North America, www.eFloras.org 26 Dec 2011

cite-note-116. citerefnrcsNRCS. "Botrychium multifidum". PLANTS Database. United States Department of Agriculture (USDA).

cite-note-127. B. multifidum University of Wisconsin, Cofrin Center for Biodiversity, Ferns and Fern Allies of Wisconsin. 26 Dec 2011

cite-note-138. BotrychiumArchived 2013-03-17 at the Wayback Machine Robert W. Freckmann Herbarium, University of Wisconsin - Stevens Point 02 Jan 2012

cite-note-149. citerefnrcsNRCS. "Botrychium oneidense". PLANTS Database. United States Department of Agriculture (USDA). Retrieved 16 Jan 2012.

cite-note-1510. B. oneidense Flora of North America, www.eFloras.org 26 Dec 2011

cite-note-1611. Botrychium oneidense Tropicos.org. Missouri Botanical Garden. 16 Jan 2012

cite-note-1712. citerefmontgomery1990Montgomery, James D. (1990). "Survivorship and Predation Changes in Five Populations of Botrychium dissectum in Eastern Pennsylvania". American Fern Journal. 4. 80 (4): 173–182. doi:10.2307/1547206. JSTOR 1547206.

cite-note-1813. B. dissectum Tropicos.org. Missouri Botanical Garden. 26 Dec 2011

cite-note-1914. B. dissectum Flora of North America, www.eFloras.org 26 Dec 2011

cite-note-2015. B. obliquum Tropicos.org. Missouri Botanical Garden. 16 Jan 2012

cite-note-2116. B. biternatum Flora of North America, www.eFloras.org 26 Dec 2011

cite-note-2222. ↑ citerefkelly1994Kelly, Dave (1994). "Demography and conservation of Botrychium australe, a peculiar, sparse mycorrhizal fern". New Zealand Journal of Botany. 32 (4): 393–400. Bibcode:1994NZJB...32..393K. doi:10.1080/0028825x.1994.10412925.

cite-note-2323. ↑ B. australe Plants for a Future 26 Dec 2011

cite-note-2424. ↑ B. biforme International Plant Names Index (IPNI) 02 Jan 2012

cite-note-2525. ↑ S. biforme International Plant Names Index (IPNI) 02 Jan 2012

cite-note-2626. ↑ B. jenmanii Flora of North America, www.eFloras.org 26 Dec 2011

cite-note-2727. ↑ B. jenmanii Tropicos.org. Missouri Botanical Garden. 26 Dec 2011

cite-note-2828. ↑ B. alabamense Tropicos.org. Missouri Botanical Garden. 27 Dec 2011

cite-note-2929. ↑ B. rugulosum Tropicos.org. Missouri Botanical Garden. 26 Dec 2011

cite-note-botrug-3030. ↑ B. rugulosum Robert W. Freckmann Herbarium, University of Wisconsin - Stevens Point 27 Dec 2011

cite-note-3131. ↑ citerefnrcsNRCS. "Sceptridium subbifoliatum". PLANTS Database. United States Department of Agriculture (USDA). Retrieved 16 Jan 2012.

cite-note-3232. ↑ B. subbifoliatum International Plant Names Index 27 Dec 2011

cite-note-3333. ↑ B. subbifoliatum ITIS taxonomy 26 Dec 2011

cite-note-3434. ↑ B. underwoodianum Tropicos.org. Missouri Botanical Garden. 27 Dec 2011

cite-note-3535. ↑ S. underwoodianum Tropicos.org. Missouri Botanical Garden. 27 Dec 2011